Likely high temperatures and/or chemical resistance, maybe very high tightening torques or force, but possibly others, too. Mostly more specialized fastening, I would guess.
But think about the bolts and nuts that hold the exhaust system to a car engine. Temperatures are going to be very high and nylon wouldn't withstand those temperatures. Things like that.
According to a NASA fastener design manual[1](page 9) split-ring style lock washers are useless for locking purposes. They say that star washers do proved some additional friction/locking action, but at the cost of damage to the surface(the friction comes from digging into the surface).
[1] https://www.google.com/search?q=nylock+nut&espv=2&source=lnm...